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Brightening theaters this weekend is Illumination Entertainment's take on Dr. Seuss's The Lorax, which features Danny DeVito as the voice of the fuzzy and colorful eco-guardian. DeVito's Lorax sports one bushy tuft of facial hair, its overgrowth stretching past the width of his waistline. The rest of cinema's most memorable mustaches can't boast that same disproportionate bulk, but they're not to be undervalued. Two are among the most iconic physical traits in film history, four make up one big whiskery package deal, and one is so indelible that its wearer spawned the name for a whole style of 'stache.

Sacha Baron Cohen

Sacha Baron Cohen in Borat (2006). Long before he was covering Ryan Seacrest in Kim Jong Il's “Bisquick ashes,” Sacha Baron Cohen was introducing the world to his ultimately inescapable Kazakh TV personality, whose lack of proper grooming was evident not just in the oversized caterpillar that rested above his lips, but in the private places laid bare thanks to one immortal neon swimsuit.


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